From the dunes on Lake Erie to the gorge-cut plateau along the Ohio River, from which Ohio takes its name, the land is fairly flat, with some pleasant rolling country and, in the southeast, small rugged hills leading to the mountains of West Virginia. Before the coming of settlers to the state, it was covered with miles of virgin forest, but today only vestiges of the trees that helped to build the many cities remain. Columbus is the capital and largest city. Cleveland is the center of the state's largest metropolitan area. Other major cities are Cincinnati, Toledo, and Akron.